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Washed ashore as a baby in tiny Tupelo Landing, North Carolina, Mo LoBeau, now eleven, and her best friend Dale turn detective when the amnesiac Colonel, owner of a cafe? and co-parent of Mo with his cook, Miss Lana, seems implicated in a murder.

Life becomes more interesting for Ananka Fishbein when, at the age of twelve, she discovers an underground room in the park across from her New York City apartment and meets a mysterious girl called Kiki Strike who claims that she, too, wants to explore the subterranean world.

Fourteen-year-olds Ananka Fishbein, Kiki Strike, and the other Irregulars encounter a Chinese mummy, a ghost, trained squirrels, and old enemies as they try to stop an art forgery ring and safeguard the secret streets hidden beneath New York City.